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The chromatophores are controlled by the nervous system and are triggered by hormones, while the initial stimulus seems to by visual. Colour change varies greatly among species, some such as the cuttlefish changing colour as the background changes. Most changes, however, appear to be behavioural, i.e. courtship or when alarmed. Many of the deep and mid-water squid are also bioluminescent, with photophores arranged in patterns over the body. This bioluminescence may be intrinsic, or it may be produced by symbiotic bacteria.