SCIENCES
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EARTH SCIENCES |
#NEXUS BEGIN DATA; DIMENSIONS NTAX=21 NCHAR=112; FORMAT SYMBOLS="0~1" MISSING=9; [CHARACTER LIST; characters 1-112 from Benton (taken from original coding sheets); 113-134 from Clark - listed in sequence from Benton & Clark (1988, p. 335 - Crocodylomropha, Unnamed group A; Unnamed group B): 1. Septomaxilla is absent; 2. Extra slit-like fenestra between the maxilla and premaxilla; 3. Movable joint between the maxilla and premaxilla; 4. Possession of antorbital fenestra; 5. Antorbital fenestra large and lies in a depression; 6. Main antorbital fenestra is low in front; 7. Nasals run forward between nares; 8. Postfrontal reduced (Evans); 9. Loss of postfrontal; 10. Tall orbit with a 'stepped' postorbital/ jugal bar behind; 11. Lacrimal forms a heavy ridge over the orbit; 12. Jugal has an anterior process between the antorbital fenestra and orbit; 13. Posterior border of lower temporal fenestra is bowed (Evans); 14. Lower temporal fenestra is reduced in size and triagnular in shape; 15. Squamosal reduced and descending ramus gracile (Gauthier); 16. Short descending process of squamosal and tall quadratojugal that contacts postorbital (Clark); 17. Postparietals are absent; 18. Postparietals fused or absent (Evans); 19. Parietal foramen absent; 20. Parietals send posterior processes into the occiput; 21. Otic notch well developed; 22. Maxillary-vomer secondary palate (Clark); 23. Pterygoids meet medially in the palate; 24. Palatal teeth are absent; 25. Marginal teeth laterally compressed and serrated; 26. Thecodont dentition; 27. Presence of an ossified laterosphenoid; 28. Pit between basioccipital and basisphenoid; 29. Enlarged pneumatic basipterygoid processes (Gauthier); 30. Lateral mandibular fenestra; 31. Coronoid reduced or absent; 32. Presacral vertebral column divided into three regions (cervical, cervical-thoracic, lumbar); 33. atlas centrum and axis intercentrum fuse; 34. axial diapophysis reduced or absent (Gauthier); 35. Diapophysis of cervicals is placed high on the neural arch; 36. Centra steeply inclined in at least cervicals 3-6 (Gauthier); 37. Presacral intercentra are absent behind the axis; 38. Cervical ribs short and stout; 39. Loss of trunk intercentra (Evans); 40. Parapophysis transfers ro neural arch in anterior dorsals; 41. Diapophysis and parapophysis fuse in posterior dorsals (i.e single-headed ribs); 42. Neural spines of dorsal vertebrae have expanded tips; 43. Dorsal centra are very constricted in ventral view; 44. Three or more sacral vertebrae; 45. Ribs are all one- or two-headed; 46. Zygapophyses of middle and posterior caudals are inclined postero-ventrally; 47. Accessory neural spine on caudals (Clark); 48. Ossified portion of scapula very tall and narrow (height more than twice width of base); 49. Scapula is long and strap-like, without expanded tip; 50. Coracoid small and glenoid faces largely backwards; 51. Glenoid faces fully backwards; 52. Loss of interlavicle; 53. Reduction or loss of clavicle; 54. Loss of clavicle (Clark); 55. Forelimb: hindlimb length ratio is c. 0.6 (bipedality); 56. Deltopectoral crest runs down one-quarter to one-third the length of he humerus; 57. Deltopecttoral crest low and runs one-third or one-half length of shaft; 58. Distal end of humerus narrower than proximal; 59. No ectepicondylar groove or foramen (Evans); 60. Manus digit I is short and equipped with a diverging claw; 61. Manus digits IV and V are reduced; 62. No more than four phalanges in manus digit IV; 63. Pelvis is markedly 3-rayed, with long downturned pubis and ischium; 64. Acetabulum perforated; 65. Acetabulum fully open (not just a slit); 66. Acetabulum subhorizontal and faces down, giving 'pillar-like' erect posture; 67. Supraacetabular crest on ilium; 68. Iliac blade is oriented subhorizontally; 69. Iliac blade is low and long; 70. Iliac blade has a small anterior process; 71. Pubis attaches to an antero-ventral face on the ilium; 72. Pubis has a strongly downturned anterior tuber, when seen in lateral view; 73. Pubis is more than three times as long as acetabular width (Gauthier); 74. Proximal distance between the ischia is less than that between the pubes; 75. No pubo-ischiadic plate (much reduced contact between pubis and ischium); 76. Pubis is long and narrow and subvertically oriented; 77. Pubis is longer than the ischium; 78. Ischium has a large postero-ventral process; 79. Hindlimbs are under the body (semi-erect or erect gait); 80. Proximal head of femur tunred inwards at about 45 degrees; 81. Proximal head of femur fully offset, with distinct neck and ball; 82. Possession of lesser trochanter; 83. Possession of fourth trochanter; 84. Fourth trochanter is a sharp flange; 85. Fourth trochanter is a wing-like process; 86. Fourth trochanter runs one-third to one-half of length of shaft; 87. Femur is ahorter than tibia; 88. Shaft of femur is bowed dorsally; 89. Distal end of femur forms two subterminal condyles; 90. Knee articulates at c. 90 degrees; 91. Prominent cnemial crest on tibia; 92. Fibula is greatly reduced; 93. Stance is digitigrade; 94. Tarsus contains only four elements 95. 'Crocodiloid' tarsus (loss of foramen; rotation between calcaneum and astragalus); 96. Crocodile-normal tarsus; 97. Advanced crocodile-normal tarsus; 98. Large posterior calcaneal tuber; 99. Ventral flange of astragalus absent (Gauthier); 100. Crocodile-reversed ankle joint (peg on calcaneum; socket on astragalus); 101. Mesotarsal ankle joint, with astragalus and calcaneum fused and to tibia; 102. Calcaneum with no tuber at all; 103. Ascending process of astragalus fits between tibia and fibula; 104. Ascending process well developed; 105. Metatarsals II, III and IV subequal, with III longest; 106. Loss of anterior proximal 'hook' on metatarsal V; 107. Digit V of the foot is reduced (shorter than I); 108. Metatarsals II-IV are closely bunched as a unit; 109. Metatarsals II-IV are elongated and foot is funtionally tridactyl; 110. Fewer than four phalanges in pedal digit V (Gauthier); 111. Dermal armour present, with two osteoderms per vertebra; 112. Osteoderms on the ventral surface of the tail; 113. Squamosal broadly ovrhangs quadrate laterally; 114. Postfrontal absent; 115. Post-temporal fenestra small or absent; 116. Prootic does not broadly contact anterior surface of paroccipital process; 117. Proximal carpals elongated; 118. Primary contact of quadrate head with prootic; 119. Pneumatic space in body of basisphenoid; 120. Mastoid antrum enters into prootic; 121. Pedal digit IV with four phalanges; 122. Quadratojugal reaches dorsally to postorbital, descending process of squamosal concomitantly reduced; 123. Parietal relatively narrow on occiput, squamosal relatively wide; 124. Maxillae form secondary palate (with vomer); 125. Jugal does not form posterior border of antorbital fenestra; 126. Quadrate foramen absent between quadrate and quadratojugal; 127. Subcapsular process on the otoccipital; 128. Posteroventral process in the coracoid; 129. Antorbital fenestra relatively small, about as tall as long; 130. Parietals fused, 131. Maxillary secondary palate without vomer contribution; 132. Lateral edge of squamosal with groove; 133. Dorsal osteoderms rectangular in shape; 134. Great anterior extent of quadrate beneath squamosal.] MATRIX Youngina 90000000000000000000000000000000990000001000001000000000000000000000000000000000000000000000000000000000000000000000000000000000000000 Trilophosaurus 00000000000000001110001100000000000000001000000100000000000000000000000000000000000000000000000000000000000000000000000000000000000000 Hyperodapedon 90000000000000001110001100000000000010111000000100000000000000000000000000000010000000000000000000000000000000000000000000000000000000 Prolacerta 90000000000000000000000000000000000000001000000000000000000000000000000000000000000000000000000000000000000000000000000000000000000000 Proterosuchus 90010001000010000100000010100000000000100000000000000000001009000000000000000000001000000000000000000000000000000000000000000000000000 Erythrosuchus 90010001000010000100000010100110000010100000000101000001011999100000010100000100001001000000010000000000110001000000000000000000000000 Euparkeria 90011011000110100110100011000110001000111100100101000001011009100000010100000110001000000000011000100000110001100000000000000000000000 Chanaresuchus 90010001100010000110110011900110000010100000100101000001011999000000010100000110001001000000011000000000111001100000000000000000000000 Parasuchus 00011001000010000111101111100110001011111100100101000000011001100000010000000110001000000000011101000000110000100000000000000000000000 Ornithosuchus 10011011000110101111101111999110991010111000100101000011011119110010010110111111011100010010111000910000111001100000000000000000000000 Doswellia 99999991109900001110191011900010990010100100100999900009999999000001110100001010001000900009999999999999999999100000000000000000000000 Stagonolepis 10011011000111011111101111110110991011111110100101000000011000100101111100111010001000000000011111000000111000110000000000000000000000 Gracilisuchus 10011011000011000111191111999110091011111100100101000000011999100000010999199111001000000010111111000000111001100000000000000000000000 Ticinosuchus 11111111011111001199199911999190111011111110101101000000011000100101111101110111001000000000011111000000111001110000000000000000000000 Saurosuchus 11111111011111901199191111999190991011111110101101099900019999100111111111110111001000000000011111000000111001190000000000000000000000 Postosuchus 11111111011101011111111111999110111011111001109101011110011000110010011110111111001000000000111111000000111001990000000000000000000000 Terrestrisuchus 90011011100001011110110111199110911011111000101101001110011001110010011011011111110010001000001111110000011100111111111110000000000000 Protosuchus 10011011100191011111111111191110111011111000101101001100011001110000010110101111001000100000011111000000111001111111111111111111111111 Crocodylus 10009091100991011111111111191110111011111000101101001100011001110000011011101011110010001000000111110000001110011111111111111111111111 Lagosuchus 90011099999999109999909911100991001110111000190101099911011999110010010110111111011111011110111000101110111111990000000000000000000000 P1ateosaurus 10011011000010101111100111100111001110111001110111111011111101111010010110111111111111111111111000101111111111100000000000000000000000 ; END;